The Wrist Speed box lets you change the speed your measuring machine will use when rotating a probe's wrist (such as a CW43L). Depending on the state of the Display absolute speeds check box in the Part/Machine tab of the Setup Options dialog box, this will either be an absolute speed (mm/sec) or a percentage of the machine's defined top speed.
The Edit window command line for this option reads:
WRISTSPEED/ nnn.nnnn